Muscle synergies inherent in simulated hypogravity running reveal flexible but not unconstrained locomotor control.

With human space exploration back in the spotlight, recent studies have investigated the neuromuscular adjustments to simulated hypogravity running. They have examined the activity of individual muscles, whereas the central nervous system may rather activate groups of functionally related muscles, known as muscle synergies. To understand how locomotor control adjusts to simulated hypogravity, we examined the temporal (motor primitives) and spatial (motor modules) components of muscle synergies in participants running sequentially at 100%, 60%, and 100% body weight on a treadmill. Our results highlighted the paradoxical nature of simulated hypogravity running: The reduced mechanical constraints allowed for a more flexible locomotor control, which correlated with the degree of spatiotemporal adjustments. Yet, the increased temporal (shortened stance phase) and sensory (deteriorated proprioceptive feedback) constraints required wider motor primitives and a higher contribution of the hamstring muscles during the stance phase. These results are a first step towards improving astronaut training protocols.

Freezing and freeing of degrees of freedom in joint action learning.

In daily life, we often encounter situations in which we have to coordinate our actions with others to achieve a common goal. These actions are also defined as joint actions. In this study we investigated how a multi-agent system learns to acquire control in a novel joint action task. To this end, we designed a task in which agents had to coordinate their actions so as to control a ball rolling on a long, hand-held beam. Participants' task was to roll the ball as fast and accurately as possible back-and-forth between two indicated targets on the beam, by manually adjusting the inclination angle of the beam. In the joint action version of this task, two participants each hold a different beam extremity. In a solo action version, the participant holds one extremity while the other is attached to a static support. The experiment consisted of two practice sessions that each comprised 15 two-min trials. One group of 12 participants first performed a solo action session of the task and then a joint action session (Group S/J), while another group of 12 participants started with a joint action session, followed by a solo action session (Group J/S). While performance increased over practice in all sessions, we found that in the joint action task dyads without prior solo task experience (Group J/S) adopted a sequential pattern of interpersonal coordination by freezing their motion whenever the other agent moved. In contrast, dyads that had received prior practice in the solo task setting (Group S/J) demonstrated less freezing and more complementary motion during the joint action performance. Lastly, we found that initial practice as a dyad in the joint action task did not result in a significant improvement of a subsequent solo action performance. We concluded that multi-agent motor learning in a novel joint action task is characterized by the initial freezing of task-relevant degrees of freedom, while individual training in a constrained setting can stimulate the freeing of these DFs during subsequent joint action performance.

How about running on Mars? Influence of sensorimotor coherence on running and spatial perception in simulated reduced gravity.

Motor control, including locomotion, strongly depends on the gravitational field. Recent developments such as lower-body positive pressure treadmills (LBPPT) have enabled studies on Earth about the effects of reduced body weight (BW) on walking and running, up to 60% BW. The present experiment was set up to further investigate adaptations to a more naturalistic simulated hypogravity, mimicking a Martian environment with additional visual information during running sessions on LBPPT. Twenty-nine participants performed three sessions of four successive five-min runs at preferred speed, alternating Earth- or simulated Mars-like gravity (100% vs. 38% BW). They were displayed visual scenes using a virtual reality headset to assess the effects of coherent visual flow while running. Running performance was characterized by normal ground reaction force and pelvic accelerations. The perceived upright and vection (visually-induced self-motion sensation)in dynamic visual environments were also investigated at the end of the different sessions. We found that BW reduction induced biomechanical adaptations independently of the visual context. Active peak force and stance time decreased, while flight time increased. Strong inter-individual differences in braking and push-off times appeared at 38% BW, which were not systematically observed in our previous studies at 80% and 60% BW. Additionally, the importance given to dynamic visual cues in the perceived upright diminished at 38% BW, suggesting an increased reliance on the egocentric body axis as a reference for verticality when the visual context is fully coherent with the previous locomotor activity. Also, while vection was found to decrease in case of a coherent visuomotor coupling at 100% BW (i.e., post-exposure influence), it remained unaffected by the visual context at 38% BW. Overall, our findings suggested that locomotor and perceptual adaptations were not similarly impacted, depending on the -simulated- gravity condition and visual context.

Neuromuscular adjustments to unweighted running: the increase in hamstring activity is sensitive to trait anxiety.

Introduction: Originally developed for astronauts, lower body positive pressure treadmills (LBPPTs) are increasingly being used in sports and clinical settings because they allow for unweighted running. However, the neuromuscular adjustments to unweighted running remain understudied. They would be limited for certain lower limb muscles and interindividually variable. This study investigated whether this might be related to familiarization and/or trait anxiety. Methods: Forty healthy male runners were divided into two equal groups with contrasting levels of trait anxiety (high, ANX+, n = 20 vs. low, ANX-, n = 20). They completed two 9-min runs on a LBPPT. Each included three consecutive 3-min conditions performed at 100%, 60% (unweighted running), and 100% body weight. Normal ground reaction force and electromyographic activity of 11 ipsilateral lower limb muscles were analyzed for the last 30 s of each condition in both runs. Results: Unweighted running showed muscle- and stretch-shortening cycle phase-dependent neuromuscular adjustments that were repeatable across both runs. Importantly, hamstring (BF, biceps femoris; STSM, semitendinosus/semimembranosus) muscle activity increased during the braking (BF: +44 ± 18%, p < 0.001) and push-off (BF: +49 ± 12% and STSM: +123 ± 14%, p < 0.001 for both) phases, and even more so for ANX+ than for ANX-. During the braking phase, only ANX+ showed significant increases in BF (+41 ± 15%, p < 0.001) and STSM (+53 ± 27%, p < 0.001) activities. During the push-off phase, ANX+ showed a more than twofold increase in STSM activity compared to ANX- (+119 ± 10% vs. +48 ± 27, p < 0.001 for both). Conclusion: The increase in hamstring activity during the braking and push-off phases may have accelerated the subsequent swing of the free-leg, likely counteracting the unweighting-induced slowing of stride frequency. This was even more pronounced in ANX+ than in ANX-, in an increased attempt not to deviate from their preferred running pattern. These results highlight the importance of individualizing LBPPT training and rehabilitation protocols, with particular attention to individuals with weak or injured hamstrings.

Timing of steering actions in locomotor interception of targets following curving trajectories.

Recent work on the visual guidance of locomotor interception of nonuniformly moving targets argued for an early reliance on first-order (velocity-based) changes in the target's bearing angle that was complemented approximately 1 second later with reliance on second-order (acceleration-based) changes. Here we provide further support for this hypothesis in a virtual driving task, in which 19 participants steered a vehicle to intercept targets moving along receding circular trajectories. Adopting a set of carefully designed target trajectories, we tested discriminating predictions with respect to the timing and direction of the first steering action. Analyses of temporal and directional characteristics of first steering events revealed a pattern of results that was fully compatible with our predictions. Moreover, application of the recently developed QuID method, focusing on the temporal co-evolution of steering behavior and the potential information sources driving it, confirmed the operative progression from early reliance on first-order changes to subsequent (after approximately 1 second) reliance on a combination of first- and second-order changes in the target's bearing angle over the course of action at the individual-trial level. The finding of an evolution over time toward higher-order informational variables, potentially captured by a fractional-order time derivative, may have consequences for other locomotor interception tasks such as running to catch a fly ball.

An Ex Situ Cadaver Liver Training Model Continuously Pressurized to Simulate Specific Skills Involved in Laparoscopic Liver Resection: the Lap-Liver Trainer.

BACKGROUND: Laparoscopic liver resection (LLR) requires delicate skills. The aim of the study was to develop a training model mimicking as much as possible intraoperative bleeding and bile leakage during LLR. We also assessed the educational value of the training model. METHODS: The Lap-liver trainer (LLT) combined a continuously pressurized ex situ cadaver liver and a customized mannequin. The customized mannequin was designed by computer-aided design and manufactured by 3D printing. The left lateral sectionectomy (LLS) was chosen to assess the feasibility of a LLR with the LLT. Eighteen volunteers were recruited to perform LLS and to assess the educational value of the LLT using a Likert scale. RESULTS: The customized mannequin consisted of a close laparoscopic training device based on a simplified reconstruction of the abdominal cavity in laparoscopic conditions. Ex situ cadaver livers were pressurized to simulate blood and bile supplies. Each expert surgeon (n = 3) performed two LLS. They were highly satisfied of simulation conditions (4.80 ± 0.45) and strongly recommended that the LLT should be incorporated into a teaching program (5.00 ± 0.0). Eight novice and 4 intermediate surgeons completed a teaching program and performed a LLS. Overall, the level of satisfaction was high (4.92 ± 0.29), and performing such a procedure under simulation conditions benefited their learning and clinical practice (4.92 ± 0.29). CONCLUSIONS: The LLT could provide better opportunities for trainees to acquire and practice LLR skills in a more realistic environment and to improve their ability to deal with specific events related to LLR.

Characterisation of visual guidance of steering to intercept targets following curving trajectories using Qualitative Inconsistency Detection.

This study explored the informational variables guiding steering behaviour in a locomotor interception task with targets moving along circular trajectories. Using a new method of analysis focussing on the temporal co-evolution of steering behaviour and the potential information sources driving it, we set out to invalidate reliance on plausible informational candidates. Applied to individual trials rather than ensemble averages, this Qualitative Inconsistency Detection (QuID) method revealed that steering behaviour was not compatible with reliance on information grounded in any type of change in the agent-centred target-heading angle. First-order changes in the environment-centred target's bearing angle could also not adequately account for the variations in behaviour observed under the different experimental conditions. Capturing the observed timing of unfolding steering behaviour ultimately required a combination of (velocity-based) first-order and (acceleration-based) second-order changes in bearing angle. While this result may point to reliance on fractional-order based changes in bearing angle, the overall importance of the present findings resides in the demonstration of the necessity to break away from the existing practice of trying to fit behaviour into a priori postulated functional strategies based on categorical differences between operative heuristic rules or control laws.

Visual guidance of locomotor interception is based on nulling changes in target bearing (not egocentric target direction nor target-heading angle).

In two experiments we studied how participants steer to intercept uniformly moving targets in a virtual driving task under hypotheses-differentiating conditions of initial target eccentricity and target motion. In line with our re-analysis of findings from earlier studies, in both experiments the observed interception behavior could not be understood as resulting from reliance on (changes in) egocentric target direction nor from reliance on (changes in) target-heading angle. The overall pattern of results observed was however compatible with a control strategy based on nulling changes in the target's bearing angle. The presence of reversals in movement direction under specific combinations of target eccentricity and motion conditions indicated that the information used was not purely rate-of-change (i.e., first-order) based but carried traces of an influence of initial target position. In Experiment 2 we explicitly tested the potential role of early reliance on perceived egocentric target direction by examining the effects of a 10° rotation of the visual scene (i.e., of both target and environment). While such a rotation gave rise to minor changes in the moment of initiation of the first steering action, contrary to predictions it did not affect the characteristics of the direction-reversal phenomenon. We conclude that the visual guidance of locomotor interception is best understood as resulting from nulling changes in the target's bearing angle, with such nulling perhaps best conceived as being fractional-order (rather integer-order) driven.

Reversals in Movement Direction in Locomotor Interception of Uniformly Moving Targets.

Here we studied how participants steer to intercept uniformly moving targets in a virtual driving task. We tested the hypothesis that locomotor interception behavior cannot fully be explained by a strategy of nulling rate of change in pertinent agent-target relations such as the target-heading angle or target's bearing angle. In line with a previously reported observation and model simulations, we found that, under specific combinations of initial target eccentricity and target motion direction, locomotor paths revealed reversals in movement direction. This phenomenon is not compatible with unique reliance on first-order (i.e., rate-of-change based) information in the case of uniformly moving targets. We also found that, as expected, such reversals in movement direction were not observed consistently over all trials of the same experimental condition: their presence depended on the timing of the first steering action effected by the participant, with only early steering actions leading to reversals in movement direction. These particular characteristics of the direction-reversal phenomenon demonstrated here for a locomotor interception-by-steering task correspond to those reported for lateral manual interception. Together, these findings suggest that control strategies operating in manual and locomotor interception may at least share certain characteristics.

Social support from evaluative familiar persons - a buffer against stress? Preliminary evidence of neuroendocrine and psychological responses.

Background and Objectives: Social support may have a stress-buffering effect when an individual is or could be negatively judged by others, but paradoxically may also exacerbate stress. The aim of our study was to examine these findings when social support was provided by a positive or negative evaluative audience composed of familiar and close others (teachers). Design and Methods: 84 men were confronted with the Trier Social Stress Test for Groups through a 3 (negative, positive, no-audience) x 2 (familiar, unfamiliar) experimental design with four measurement points of cortisol levels and state anxiety. We also tested whether closeness with the committee members predicted these variables for the participants in the familiar conditions. Results: Using both a frequentist and a Bayesian approach, familiarity and social support did not have stress-buffering effects (or merely anecdotal effects) on cortisol levels but buffered self-reported anxiety only for the participants faced with a supportive audience composed of familiar persons. Closeness with the experimenters was not a significant predictor of the stress responses. Conclusions: Because these results are preliminary evidence, further investigations into the relations between support provider and recipient during evaluative tasks would be worthwhile to better explain opposing findings found in this growing literature.

Information-Based Social Coordination Between Players of Different Skill in Doubles Pong.

We studied how teams of two players of different skill level intercepted approaching balls in the doubles-pong task. In this task, the two players moved their on-screen paddles along a shared interception axis, so that the approaching ball was intercepted by one of the paddles and that the paddles did not collide. Earlier work revealed the presence of a fuzzy division of interception space, with a boundary between interception domains located in the space between the two initial paddle positions. In the present study, using the performance of the players in their individual training sessions, we formed teams of players of varying skill level. We considered two accounts of how this boundary should be understood. In a first account, the players have shared knowledge of this boundary. Based on the side of the boundary at which the approaching ball will cross the interception axis, the players would decide whose paddle is to make the interception. Under this account, we expected that a better-skilled player would take responsibility for a larger interception domain, leading to a boundary closer to the lesser-skilled player. However, our analyses did not reveal any systematic effect of skill difference on the location (or degree of fuzziness) of the boundary: location of boundaries and overlap of interception domains varied over teams but were not systematically related to skill differences between team members. We did find effects of ball speed and approach angle. In a second account, the boundary emerges from (information-driven) player-player-ball interactions. An action-based model consistent with this account was able to capture all the patterns in boundary positions and overlaps that we observed. We conclude that the interception patterns that players demonstrate in the doubles-pong task are best understood as emerging from the unfolding of the dynamics of the system of the two players and the ball, coupled through information.

Division of labor as an emergent phenomenon of social coordination: The example of playing doubles-pong.

In many daily situations, our behavior is coordinated with that of others. This study investigated this coordination in a doubles-pong task. In this task, two participants each controlled a paddle that could move laterally near the bottom of a shared computer screen. With their paddles, the players needed to block balls that moved down under an angle. In doing so, they needed to make sure that their paddles did not collide. A successful interception led to the ball bouncing back upwards. Importantly, all communication other than through vision of the shared screen was excluded. In the experiment, the initial position of the paddle of the right player was varied across trials. This allowed testing hypotheses regarding the use of a tacitly understood boundary to divide interception space. This boundary could be halfway the screen, or in the middle between the initial positions of the two paddles. These two hypotheses did not hold. As an alternative to planned division of labor, the behavioral patterns might emerge from continuous visual couplings of paddles and ball. This was tested with an action-based decision model that considered the rates of change of each player's angle between the interception axis and the line connecting the ball and inner edge of the paddle. The model accounted for the observed patterns of behavior to a very large extent. This led to the conclusion that decisions of who would take the ball emerged from ongoing social coordination. Implications for social coordination in general are discussed.

Playing 'Pong' Together: Emergent Coordination in a Doubles Interception Task.

In this contribution we set out to study how a team of two players coordinated their actions so as to intercept an approaching ball. Adopting a doubles-pong task, six teams of two participants each intercepted balls moving downward across a screen toward an interception axis by laterally displacing participant-controlled on-screen paddles. With collisions between paddles resulting in unsuccessful interception, on each trial participants had to decide amongst them who would intercept the ball and who would not. In the absence of possibilities for overt communication, such team decisions were informed exclusively by the visual information provided on the screen. Results demonstrated that collisions were rare and that 91.3 ± 3.4% of all balls were intercepted. While all teams demonstrated a global division of interception space, boundaries between interception domains were fuzzy and could moreover be shifted away from the center of the screen. Balls arriving between the participants' initial paddle positions often gave rise to both participants initiating an interception movement, requiring one of the participants to abandon the interception attempt at some point so as to allow the other participant to intercept the ball. A simulation of on-the-fly decision making of who intercepted the ball based on a measure capturing the triangular relations between the two paddles and the ball allowed the qualitative aspects of the pattern of observed results to be reproduced, including the timing of abandoning. Overall, the results thus suggest that decisions regarding who intercepts the ball emerge from between-participant interactions.

The Influence of the 'Trier Social Stress Test' on Free Throw Performance in Basketball: An Interdisciplinary Study.

The aim of the present study was to explore the relationship between stress and sport performance in a controlled setting. The experimental protocol used to induce stress in a basketball free throw was the Trier Social Stress Test (TSST) and its control condition (Placebo-TSST). Participants (n = 19), novice basketball players but trained sportspersons, were exposed to two counterbalanced conditions in a crossover design. They were equipped with sensors to measure movement execution, while salivary cortisol and psychological state were also measured. The task consisted of two sequences of 40 free throws, one before either the TSST or Placebo-TSST and one after. Physiological and psychological measures evidenced that the TSST induced significant stress responses, whereas the Placebo-TSST did not. Shooting performance remained stable after the TSST but decreased after the Placebo-TSST. We found no effect of the TSST or Placebo-TSST on movement execution. A multivariate model of free throw performance demonstrated that timing, smoothness and explosiveness of the movements are more relevant to account for beginner's behavior than stress-related physiological and psychological states. We conclude that the TSST is a suitable protocol to induce stress responses in sport context, even though the effects on beginners' free throw performance and execution are small and complex.

Fractional-order information in the visual control of lateral locomotor interception.

Previous work on locomotor interception of a target moving in the transverse plane has suggested that interception is achieved by maintaining the target's bearing angle (often inadvertently confused and/or confounded with the target heading angle) at a constant value. However, dynamics-based model simulations testing the veracity of the underlying control strategy of nulling the rate of change in the bearing angle have been restricted to limited conditions of target motion, and only a few alternatives have been considered. Exploring a wide range of target motion characteristics with straight and curving ball trajectories in a virtual reality setting, we examined how soccer goalkeepers moved along the goal line to intercept long-range shots on goal, a situation in which interception is naturally constrained to movement along a single dimension. Analyses of the movement patterns suggested reliance on combinations of optical position and velocity for straight trajectories and optical velocity and acceleration for curving trajectories. As an alternative to combining such standard integer-order derivatives, we demonstrate with a simple dynamical model that nulling a single informational variable of a self-tuned fractional (rather than integer) order efficiently captures the timing and patterning of the observed interception behaviors. This new perspective could fundamentally change the conception of what perceptual systems may actually provide, both in humans and in other animals. (PsycINFO Database Record

Reading from a Head-Fixed Display during Walking: Adverse Effects of Gaze Stabilization Mechanisms.

Reading performance during standing and walking was assessed for information presented on earth-fixed and head-fixed displays by determining the minimal duration during which a numerical time stimulus needed to be presented for 50% correct naming answers. Reading from the earth-fixed display was comparable during standing and walking, with optimal performance being attained for visual character sizes in the range of 0.2° to 1°. Reading from the head-fixed display was impaired for small (0.2-0.3°) and large (5°) visual character sizes, especially during walking. Analysis of head and eye movements demonstrated that retinal slip was larger during walking than during standing, but remained within the functional acuity range when reading from the earth-fixed display. The detrimental effects on performance of reading from the head-fixed display during walking could be attributed to loss of acuity resulting from large retinal slip. Because walking activated the angular vestibulo-ocular reflex, the resulting compensatory eye movements acted to stabilize gaze on the information presented on the earth-fixed display but destabilized gaze from the information presented on the head-fixed display. We conclude that the gaze stabilization mechanisms that normally allow visual performance to be maintained during physical activity adversely affect reading performance when the information is presented on a display attached to the head.

Perception of spin and the interception of curved football trajectories.

Using plain white and chequered footballs, we evaluated observers' sensitivity to rotation direction and the effects of ball texture on interceptive behaviour. Experiment 1 demonstrated that the maximal distance at which observers (n = 8) could perceive the direction of ball rotation decreased when rotation frequency increased from 5 to 11 Hz. Detection threshold distances were nevertheless always larger for the chequered (decreasing from 47 to 28 m) than for the white (decreasing from 15 to 11 m) ball. In Experiment 2, participants (n = 7) moved laterally along a goal line to intercept the two balls launched with or without ±4.3 Hz sidespin from a 30-m distance. The chequered ball gave rise to shorter movement initiation times when trajectories curved outward (±6 m arrival positions) or did not curve (±2 m arrival positions). Inward curving trajectories, arriving at the same ±2 m distances from the participants as the non-curving trajectories, evoked initial movements in the wrong direction for both ball types, but the amplitude and duration of these reversal movements were attenuated for the chequered ball. We conclude that the early detection of rotation permitted by the chequered ball allowed modulation of interception behaviour without changing its qualitative characteristics.

Prospective control in catching: the persistent Angle-of-approach effect in lateral interception.

In lateral interception tasks balls converging onto the same interception location via different trajectories give rise to systematic differences in the kinematics of hand movement. While it is generally accepted that this angle-of-approach effect reflects the prospective (on-line) control of movement, controversy exists with respect to the information used to guide the hand to the future interception location. Based on the pattern of errors observed in a task requiring visual extrapolation of line segments to their intersection with a second line, angle-of-approach effects in lateral interception have been argued to result from perceptual biases in the detection of information about the ball's future passing distance along the axis of hand movement. Here we demonstrate that this account does not hold under experimental scrutiny: The angle-of-approach effect still emerged when participants intercepted balls moving along trajectories characterized by a zero perceptual bias with respect to the ball's future arrival position (Experiment 4). Designing and validating such bias-controlled trajectories were done using the line-intersection extrapolation task (Experiments 2 and 3). The experimental set-up used in the present series of experiments was first validated for the lateral interception and the line-intersection extrapolation tasks: In Experiment 1 we used rectilinear ball trajectories to replicate the angle-of-approach effect in lateral interception of virtual balls. Using line segments extracted from these rectilinear ball trajectories, in Experiment 2 we replicated the reported pattern of errors in the estimated locus of intersection with the axis of hand movement. We used these errors to develop a set of bias-free trajectories. Experiment 3 confirmed that the perceptual biases had been corrected for successfully. We discuss the implications on the information-based regulation of hand movement of our finding that the angle-of-approach effect in lateral interception cannot not explained by perceptual biases in information about the ball's future passing distance.

Temporal evolution of skeletal regenerated tissue: what can mechanical investigation add to biological?

The objective here was to experimentally characterize the temporal evolution of the structural and mechanical properties of large volume immature regenerated tissues. We studied these evolving tissues from their genesis in controlled mechanical conditions. We developed an animal model based on the periosteal properties leading to unloaded regenerated skeletal tissue. To characterize the temporal evolution of mechanical properties, we carried out indentation tests coupled with macroscopic examinations and histological studies. This combined methodology yielded a range of information on osteogenesis at different scales: macroscopic by simple observation, mesoscopic by indentation test and microscopic by histological study. Results allowed us to identify different periods, providing a link between biological changes and material property evolution in bone tissue regeneration. The regenerated tissue evolves from a viscous, homogeneous, soft material to a heterogeneous stiffer material endowed with a lower viscosity. From a biological point of view, cell organization progresses from a proliferated cell clot to a mature structure closer to that of the bone. During the first 7 days, mechanical and biological results revealed the same evolution: first, the regenerated tissue grew, then, differentiated into an osteochondral tissue and finally calcification began. While our biological results confirm those of other studies, our mechanical results provide the first experimental mechanical characterization by reduced Young's modulus of such tissue.

A kinematic analysis of rugby lineout throwing.

To characterize rugby union lineout throwing technique, three experienced male rugby players performed throwing trials under varying conditions of distance and trajectory. Motion analysis permitted the recovery of joint centre coordinates at 120 Hz and the construction of a three-dimensional linked segment model for calculation of joint angle and centre of mass time histories. All participants exhibited greater accuracy at shorter throwing distances, although the accuracy decrement was less in players of higher standard. Participants demonstrated different alterations in technique when performing throws of longer distances, either showing increased magnitudes of upper-body joint angle velocities (less accurate thrower) or lower-body joint velocities (more accurate thrower). The most elite thrower exhibited greater consistency in timing of peak joint angle velocities, with an overall standard deviation of 0.008 s compared with 0.027 s for the least accurate thrower. Data from participants of lesser ability suggest that changes are made to both magnitudes and timing of joint kinematics, which leads to increased variability in performance. The implications for players and coaches include the need to develop core strength to permit limited changes to the timing and magnitude of upper-body joint actions while allowing sufficient end-point velocity to be imparted on the ball.